The other side of Mega Man Legends! Now Mega Man's female rival, Tron from Mega Man Legends is back with her trusty Servbots in an entirely new adventure! Join Tron and the Servbots as they circle the world in their air ship, Gesellshaft, in search of adventure and priceless treasure. Brave Tron's wild exploits in q unique mix of action-adventure, RPG, and puzzles in this irresistible game for all ages. Unique gameplay and storyline, mixing the best action-adventure, RPG, and puzzles in a vast 3-D world. Vibrant colorful graphics! -explore huge 3-D environments for puzzles, treasures and other surprises "Guide Shot System" sends Servbots to retrieve treasure and perform Tron's bidding Includes Demo of Mega Man Legends 2

The game begins some time before the events of Mega Man Legends. The Bonne family's leader, Teisel, seeks the ancient Nakkai Ruins to try and uncover Diana's Tear, a gigantic and highly valuable Refractor. After rendezvousing with younger brother Bon, the two are subdued and captured by Glyde, a rival air pirate in the service of loan shark Lex Loath. Tron finds out that the money used to build the Bonnes' flying fortress was funded by Loath, and can only watch as her brothers are taken away. After discovering that Teisel defaulted on his 1,000,000-zenny loan, she realizes that she must pay back the money or Teisel and Bon will not be seen again. Seeing no other options, Tron suits up in a custom Gustaff mecha and, along with her army of 40 identical Servbots, begins her quest to pay the ransom through any means possible. During repeated bank robberies, Tron encounters officer Denise Marmalade, who repeatedly fails to stop her, even when engaging Tron in her own police-issue mecha. In the caves, the Servbots find the three Aurora Stones, help the restless spirit of a man that died trying to find the Fountain of Youth, and help two Diggers fall in love. In the ruins, Tron explores for valuable artifacts, including the Diana's Tear. After various adventures, Tron brings the one million zenny to Loath, who then claims that Teisel also owes an additional two million zenny of interest on his loan. While she manages to acquire the requested money, Loath insists that she owes interest on the interest. Realizing that Loath will never let her brothers go, she too is captured and imprisoned. Tron's favorite Servbot takes command in her absence, leading the others on a rescue mission. Though the Servbots succeed in freeing the Bonnes and Tron defeats Glyde, Loath activates his secret weapon, the Colossus. The Bonnes at first try to attack it with the Gesselschaft's weapons, but this has almost no effect, and the Colossus' return fire causes Tron to be very seriously injured and Teisel to get thrown overboard. The Servbot crew band together and defeat to take on the Colossus in the Gustaff. They succeed, finally defeating Loath and Glyde once and for all. To ensure they never cause trouble again, Tron hands Loath and Glyde into the police, saving officer Denise's job in the process. The issue finally resolved, the Bonnes, reunited, set off for Kattelox Island. While in flight, the favorite Servbot accidentally throws out a giant Refractor won from Loath with the trash, causing Tron and Teisel to panic and force a pit stop to search for it.